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Structure of paused transcription complex Pol II-DSIF-NELF - tilted
Resolution: 3.5 Å
EM Method: Single-particle
Fitted PDBs: 8uha
Q-score: 0.28
Mol Cell (2024) [ DOI: doi:10.1016/j.molcel.2024.01.023 Pubmed: 38401543 ]
- Dna-directed rna polymerase subunit beta (134 kDa, Protein from Sus scrofa)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb3 (30 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Rna polymerase rpb4/rpc9 core domain-containing protein (16 kDa, Protein from Sus scrofa)
- Transcription elongation factor spt4 (13 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Sus scrofa)
- Negative elongation factor a (57 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Sus scrofa)
- Rna polymerase in complex with dsif and nelf (Complex)
- Dna-directed rna polymerase subunit (217 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit j (13 kDa, Protein from Sus scrofa)
- Dna (28-mer) (15 kDa, DNA from synthetic construct)
- Negative elongation factor e (43 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Sus scrofa)
- Dna (38-mer) (15 kDa, DNA from synthetic construct)
- Rna polymerase ii subunit k (7 kDa, Protein from Sus scrofa)
- Rna (5 kDa, RNA from synthetic construct)
- Negative elongation factor c/d (66 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Negative elongation factor b (65 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Sus scrofa)
Structure of paused transcription complex Pol II-DSIF-NELF - post-translocated
Resolution: 2.8 Å
EM Method: Single-particle
Fitted PDBs: 8uhd
Q-score: 0.353
Mol Cell (2024) [ DOI: doi:10.1016/j.molcel.2024.01.023 Pubmed: 38401543 ]
- Dna-directed rna polymerase subunit beta (134 kDa, Protein from Sus scrofa)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Non-template dna (15 kDa, DNA from synthetic construct)
- Dna-directed rna polymerase ii subunit rpb3 (30 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Rna polymerase rpb4/rpc9 core domain-containing protein (16 kDa, Protein from Sus scrofa)
- Template dna (15 kDa, DNA from synthetic construct)
- Transcription elongation factor spt4 (13 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Sus scrofa)
- Negative elongation factor a (57 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Sus scrofa)
- Rna polymerase in complex with dsif and nelf (Complex)
- Dna-directed rna polymerase subunit (217 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit j (13 kDa, Protein from Sus scrofa)
- Negative elongation factor e (43 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit k (7 kDa, Protein from Sus scrofa)
- Rna (6 kDa, RNA from synthetic construct)
- Negative elongation factor c/d (66 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Negative elongation factor b (65 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Sus scrofa)
Structure of transcription complex Pol II-DSIF-NELF-TFIIS
Resolution: 3.23 Å
EM Method: Single-particle
Fitted PDBs: 8uis
Q-score: 0.319
Mol Cell (2024) 84 pp. 1243- [ DOI: doi:10.1016/j.molcel.2024.01.023 Pubmed: 38401543 ]
- Non-template dna (15 kDa, DNA from synthetic construct)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase subunit beta (134 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerase ii subunit rpb3 (30 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Rna polymerase rpb4/rpc9 core domain-containing protein (16 kDa, Protein from Sus scrofa)
- Template dna (15 kDa, DNA from synthetic construct)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Sus scrofa)
- Negative elongation factor a (57 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerase subunit (217 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit j (13 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit k (7 kDa, Protein from Sus scrofa)
- Rna (6 kDa, RNA from synthetic construct)
- Negative elongation factor c/d (66 kDa, Protein from Homo sapiens)
- Transcription elongation factor a protein 1 (34 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Rna polymerase in complex with dsif, nelf, tfiis (Complex)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Sus scrofa)
Structure of mammalian Pol II-TFIIS elongation complex
Resolution: 3.0 Å
EM Method: Single-particle
Fitted PDBs: 8a40
Q-score: 0.486
Farnung L, Ochmann M, Garg G, Vos SM, Cramer P
Mol Cell (2022) 82 pp. 3126-3134.e7 [ Pubmed: 35858621 DOI: doi:10.1016/j.molcel.2022.06.027 ]
- Dna-directed rna polymerase ii subunit rpb11-a (13 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit k (7 kDa, Protein from Sus scrofa)
- Non-template dna (14 kDa, DNA from synthetic construct)
- Rna polymerase ii subunit d (16 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Sus scrofa)
- Rna (4 kDa, RNA from synthetic construct)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from Sus scrofa)
- Mammalian pol ii-tfiis elongation complex (Complex from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from Sus scrofa)
- Transcription elongation factor a protein 1 (34 kDa, Protein from Homo sapiens)
- Template dna (12 kDa, DNA from synthetic construct)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerase subunit (218 kDa, Protein from Sus scrofa)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from Sus scrofa)
Pol II-DSIF-SPT6-PAF1c-TFIIS complex with rewrapped nucleosome
Resolution: 3.0 Å
EM Method: Single-particle
Fitted PDBs: 7unc
Q-score: 0.344
Filipovski M, Soffers JHM, Vos SM, Farnung L
Science (2022) 376 pp. 1313-1316 [ Pubmed: 35709268 DOI: doi:10.1126/science.abo3851 ]
- Rna polymerase ii subunit k (7 kDa, Protein from pig)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from pig)
- Transcription elongation factor spt6 (199 kDa, Protein from Homo sapiens)
- Histone h4 (11 kDa, Protein from Xenopus laevis)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from pig)
- Rpb10 (7 kDa, Protein from Sus scrofa)
- Rpb8 (17 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Histone h2a (14 kDa, Protein from Xenopus laevis)
- Rpol4c domain-containing protein (20 kDa, Protein from pig)
- Rna polymerase ii-associated factor 1 homolog (60 kDa, Protein from Homo sapiens)
- Wdr61 (33 kDa, Protein from Homo sapiens)
- Pol ii-dsif-spt6-paf1c-tfiis complex with rewrapped nucleosome (Complex)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from pig)
- Rna polymerase-associated protein leo1 (75 kDa, Protein from Homo sapiens)
- Non-template dna (64 kDa, DNA from synthetic construct)
- Transcription elongation factor a protein 1 (34 kDa, Protein from Homo sapiens)
- Histone h3.2 (15 kDa, Protein from Xenopus laevis)
- Rna (5 kDa, RNA from synthetic construct)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from pig)
- Parafibromin (60 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from pig)
- Rna polymerase-associated protein rtf1 homolog (80 kDa, Protein from Homo sapiens)
- Rpb11 (13 kDa, Protein from Sus scrofa)
- Template dna (66 kDa, DNA from synthetic construct)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Histone h2b 1.1 (13 kDa, Protein from Xenopus laevis)
- Dna-directed rna polymerase subunit (219 kDa, Protein from pig)
- Rpb9 (14 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Rna polymerase-associated protein ctr9 homolog (134 kDa, Protein from Homo sapiens)
Pol II-DSIF-SPT6-PAF1c-TFIIS-nucleosome complex (stalled at +38)
Resolution: 3.0 Å
EM Method: Single-particle
Fitted PDBs: 7und
Q-score: 0.216
Filipovski M, Soffers JHM, Vos SM, Farnung L
Science (2022) 376 pp. 1313-1316 [ Pubmed: 35709268 DOI: doi:10.1126/science.abo3851 ]
- Rna polymerase ii subunit k (7 kDa, Protein from pig)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from pig)
- Transcription elongation factor spt6 (199 kDa, Protein from Homo sapiens)
- Histone h4 (11 kDa, Protein from Xenopus laevis)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from pig)
- Rpb10 (7 kDa, Protein from Sus scrofa)
- Rpb8 (17 kDa, Protein from Sus scrofa)
- Zinc ion (65 Da, Ligand)
- Histone h2a (14 kDa, Protein from Xenopus laevis)
- Rpol4c domain-containing protein (20 kDa, Protein from pig)
- Rna polymerase ii-associated factor 1 homolog (60 kDa, Protein from Homo sapiens)
- Wdr61 (33 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc2 (14 kDa, Protein from pig)
- Transcription elongation factor spt4 (13 kDa, Protein from Homo sapiens)
- Rna polymerase-associated protein leo1 (75 kDa, Protein from Homo sapiens)
- Rna (5 kDa, RNA from synthetic construct)
- Non-template dna (64 kDa, DNA from synthetic construct)
- Transcription elongation factor a protein 1 (34 kDa, Protein from Homo sapiens)
- Histone h3.2 (15 kDa, Protein from Xenopus laevis)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from pig)
- Parafibromin (60 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from pig)
- Rna polymerase-associated protein rtf1 homolog (80 kDa, Protein from Homo sapiens)
- Pol ii-dsif-spt6-paf1c-tfiis complex (stalled at +38) (Complex)
- Rpb11 (13 kDa, Protein from Sus scrofa)
- Template dna (66 kDa, DNA from synthetic construct)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Histone h2b 1.1 (13 kDa, Protein from Xenopus laevis)
- Dna-directed rna polymerase subunit (219 kDa, Protein from pig)
- Rpb9 (14 kDa, Protein from Sus scrofa)
- Magnesium ion (24 Da, Ligand)
- Rna polymerase-associated protein ctr9 homolog (134 kDa, Protein from Homo sapiens)
Resolution: 3.3 Å
EM Method: Single-particle
Fitted PDBs: 7okx
Q-score: 0.571
Chen Y, Vos SM, Dienemann C, Ninov M, Urlaub H, Cramer P
Mol Cell (2021) 81 pp. 3386-3399.e10 [ Pubmed: 34265249 DOI: doi:10.1016/j.molcel.2021.06.019 ]
- Dna and rna (Complex from HIV whole-genome vector AA1305#18)
- Transcription elongation factor spt5 (Complex from Homo sapiens)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from Pig)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Zinc ion (65 Da, Ligand)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Pig)
- Transcription elongation complex of rna polymerase ii with elongation factors dsif(spt5-kow5)and ell2-eaf1 (Complex)
- Rna_pol_l_2 domain-containing protein (13 kDa, Protein from Pig)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Pig)
- Magnesium ion (24 Da, Ligand)
- Dna-directed rna polymerase ii subunit f (14 kDa, Protein from Pig)
- Rna polymerase ii subunit k (7 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Pig)
- Rna polymerase ii subunit d (20 kDa, Protein from Pig)
- Non-template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Rna polymerase ii elongation factor ell2 (72 kDa, Protein from Homo sapiens)
- Template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Rna (15 kDa, RNA from HIV whole-genome vector AA1305#18)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Pig)
- Ell-associated factor 1 (29 kDa, Protein from Homo sapiens)
- Elongation and association factors (Complex from Homo sapiens)
- Rna polymerase (Complex from Sus scrofa)
- Dna-directed rna polymerase ii subunit rpb1 (217 kDa, Protein from Pig)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Pig)
Structure of active transcription elongation complex Pol II-DSIF-ELL2-EAF1, composite map 2
Resolution: 4.14 Å
EM Method: Single-particle
Fitted PDBs: 7oky
Q-score: 0.387
Chen Y, Vos SM, Dienemann C, Ninov M, Urlaub H, Cramer P
Mol Cell (2021) 81 pp. 3386-3399.e10 [ Pubmed: 34265249 DOI: doi:10.1016/j.molcel.2021.06.019 ]
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from Pig)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Zinc ion (65 Da, Ligand)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Pig)
- Rna polymerase (Complex from Sus scrofa)
- Elongation factor (Complex from Homo sapiens)
- Transcription elongation factor spt4 (13 kDa, Protein from Homo sapiens)
- Rna_pol_l_2 domain-containing protein (13 kDa, Protein from Pig)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from Pig)
- Transcription elongation complex of rna polymerase ii with elongation factors dsif and ell2, eaf1 (Complex)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Pig)
- Magnesium ion (24 Da, Ligand)
- Dna-directed rna polymerase ii subunit f (14 kDa, Protein from Pig)
- Rna polymerase ii subunit k (7 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Pig)
- Non-template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Rna polymerase ii subunit d (20 kDa, Protein from Pig)
- Rna polymerase ii elongation factor ell2 (72 kDa, Protein from Homo sapiens)
- Template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Rna (15 kDa, RNA from HIV whole-genome vector AA1305#18)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Pig)
- Ell-associated factor 1 (29 kDa, Protein from Homo sapiens)
- Rna and dna (Complex from HIV whole-genome vector AA1305#18)
- Dna-directed rna polymerase ii subunit rpb1 (217 kDa, Protein from Pig)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Pig)
- Elongation and association factor (Complex from Homo sapiens)
Resolution: 3.0 Å
EM Method: Single-particle
Fitted PDBs: 7ol0
Q-score: 0.558
Chen Y, Vos SM, Dienemann C, Ninov M, Urlaub H, Cramer P
Mol Cell (2021) 81 pp. 3386-3399.e10 [ Pubmed: 34265249 DOI: doi:10.1016/j.molcel.2021.06.019 ]
- Transcription elongation complex of rna polymerase ii with elongation factors dsif (spt5-kow5) (Complex from Sus scrofa)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from Pig)
- Transcription elongation factor spt5 (121 kDa, Protein from Homo sapiens)
- Zinc ion (65 Da, Ligand)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Pig)
- Rna_pol_l_2 domain-containing protein (13 kDa, Protein from Pig)
- Dna-directed rna polymerase subunit beta (142 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit f (14 kDa, Protein from Pig)
- Rna polymerase ii subunit k (7 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Pig)
- Dna-directed rna polymerase ii subunit rpb1 (217 kDa, Protein from Sus scrofa)
- Rna polymerase ii subunit d (20 kDa, Protein from Pig)
- Non-template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Template dna (14 kDa, DNA from HIV whole-genome vector AA1305#18)
- Rna (15 kDa, RNA from HIV whole-genome vector AA1305#18)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Pig)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Pig)
- Magnesium ion (24 Da, Ligand)
Overall map of a transcribing RNA polymerase II-U1 snRNP complex with 145-nt RNA
Resolution: 3.6 Å
EM Method: Single-particle
Fitted PDBs: 7b0y
Q-score: 0.344
Zhang S, Aibara S, Vos SM, Agafonov DE, Luhrmann R, Cramer P
Science (2021) 371 pp. 305-309 [ DOI: doi:10.1126/science.abf1870 Pubmed: 33446560 ]
- Small nuclear ribonucleoprotein g (8 kDa, Protein from Homo sapiens)
- U1 small nuclear ribonucleoprotein a (31 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein-associated protein (23 kDa, Protein from Homo sapiens)
- U1 snrna (52 kDa, RNA from Homo sapiens)
- Small nuclear ribonucleoprotein sm d3 (13 kDa, Protein from Homo sapiens)
- The transcribing rna polymerase ii-u1 snrnp complex with 145-nt rna (Complex)
- Zinc ion (65 Da, Ligand)
- Small nuclear ribonucleoprotein f (9 kDa, Protein from Homo sapiens)
- U1 snrnp (Complex from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc5 (7 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit rpb3 (31 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit d (16 kDa, Protein from Sus scrofa domesticus)
- Small nuclear ribonucleoprotein sm d2 (13 kDa, Protein from Homo sapiens)
- Dna-directed rna polymerases i, ii, and iii subunit rpabc3 (17 kDa, Protein from Sus scrofa domesticus)
- 145-nt rna (46 kDa, RNA from synthetic construct)
- Magnesium ion (24 Da, Ligand)
- Dna-directed rna polymerase ii subunit rpb9 (14 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit rpb1 (217 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit rpb7 (19 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit e (24 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase subunit beta (134 kDa, Protein from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit f (14 kDa, Protein from Sus scrofa domesticus)
- Rna polymerase ii (Complex from Sus scrofa domesticus)
- Dna-directed rna polymerase ii subunit rpb11-a (13 kDa, Protein from Sus scrofa domesticus)
- U1 small nuclear ribonucleoprotein 70 kda (51 kDa, Protein from Homo sapiens)
- Nucleic acids (Complex from synthetic construct)
- Template dna (14 kDa, DNA from synthetic construct)
- Small nuclear ribonucleoprotein e (10 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein sm d1 (13 kDa, Protein from Homo sapiens)
- Non-template dna (14 kDa, DNA from synthetic construct)
- Rpb12 (7 kDa, Protein from Sus scrofa domesticus)