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Crystal Structure of bovine Arp2/3 complex

X-ray diffraction
2.65Å resolution
PDB entry 2p9l coloured by chain, front view.
PDB entry 2p9l coloured by chain, side view.
PDB entry 2p9l coloured by chain, top view.
Source organism: Bos taurus
Assembly composition:
hetero heptamer (preferred)
Entry contents: 7 distinct polypeptide molecules

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Assemblies

Assembly 1 (preferred)
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Multimeric state: hetero heptamer
Accessible surface area: 67959.39 Å2
Buried surface area: 19673.1 Å2
Dissociation area: 899.22 Å2
Dissociation energy (ΔGdiss): 0.74 kcal/mol
Dissociation entropy (TΔSdiss): 13.21 kcal/mol
Symmetry number: 1
PDBe Complex ID: PDB-CPX-108370

Macromolecules

Actin-related protein 3
Chain: A
Length: 418 amino acids
Theoretical weight: 47.43 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: P61157 (Residues: 1-418; Coverage: 100%)
Gene names: ACTR3, ARP3
Pfam: Actin
InterPro:
CATH:
SCOP: Actin/HSP70
Actin-related protein 3 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 3 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 3 in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2
Chain: B
Length: 394 amino acids
Theoretical weight: 44.82 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: A7MB62 (Residues: 1-394; Coverage: 100%)
Gene names: ACTR2, ARP2
Pfam: Actin
InterPro:
CATH:
Actin-related protein 2 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2 in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2/3 complex subunit 1B
Chain: C
Length: 372 amino acids
Theoretical weight: 41.02 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: Q58CQ2 (Residues: 1-372; Coverage: 100%)
Gene name: ARPC1B
Pfam: WD domain, G-beta repeat
InterPro:
CATH: YVTN repeat-like/Quinoprotein amine dehydrogenase
SCOP: WD40-repeat
Actin-related protein 2/3 complex subunit 1B in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2/3 complex subunit 1B in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2/3 complex subunit 1B in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2/3 complex subunit 2
Chain: D
Length: 300 amino acids
Theoretical weight: 34.4 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: Q3MHR7 (Residues: 1-300; Coverage: 100%)
Gene name: ARPC2
Pfam: Arp2/3 complex, 34 kD subunit p34-Arc
InterPro:
CATH: Yope Regulator; Chain: A,
SCOP: Arp2/3 complex subunits
Actin-related protein 2/3 complex subunit 2 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2/3 complex subunit 2 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2/3 complex subunit 2 in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2/3 complex subunit 3
Chain: E
Length: 178 amino acids
Theoretical weight: 20.57 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: Q3T035 (Residues: 1-178; Coverage: 100%)
Gene name: ARPC3
Pfam: ARP2/3 complex ARPC3 (21 kDa) subunit
InterPro:
CATH: Actin-related protein 2/3 complex subunit 3
SCOP: Arp2/3 complex 21 kDa subunit ARPC3
Actin-related protein 2/3 complex subunit 3 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2/3 complex subunit 3 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2/3 complex subunit 3 in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2/3 complex subunit 4
Chain: F
Length: 168 amino acids
Theoretical weight: 19.7 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: Q148J6 (Residues: 1-168; Coverage: 100%)
Gene name: ARPC4
Pfam: ARP2/3 complex 20 kDa subunit (ARPC4)
InterPro:
CATH: Yope Regulator; Chain: A,
SCOP: Arp2/3 complex subunits
Actin-related protein 2/3 complex subunit 4 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2/3 complex subunit 4 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2/3 complex subunit 4 in PDB entry 2p9l, assembly 1, top view.
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Actin-related protein 2/3 complex subunit 5
Chain: G
Length: 151 amino acids
Theoretical weight: 16.3 KDa
Source organism: Bos taurus
Expression system: Not provided
UniProt:
  • Canonical: Q3SYX9 (Residues: 1-151; Coverage: 100%)
Gene name: ARPC5
Pfam: ARP2/3 complex 16 kDa subunit (p16-Arc)
InterPro:
CATH: Actin-related protein 2/3 complex subunit 5
SCOP: Arp2/3 complex 16 kDa subunit ARPC5
Actin-related protein 2/3 complex subunit 5 in PDB entry 2p9l, assembly 1, front view.
Actin-related protein 2/3 complex subunit 5 in PDB entry 2p9l, assembly 1, side view.
Actin-related protein 2/3 complex subunit 5 in PDB entry 2p9l, assembly 1, top view.
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