q towards e2 if and only if q is a relational quality such that q inheres-in some e, and e != e2 and q is dependent on e2

http://www.ncbi.nlm.nih.gov/pubmed/20064205

This relation is provided in order to support the use of relational qualities such as 'concentration of'; for example, the concentration of C in V is a quality that inheres in V, but pertains to C.

Chris Mungall