EC - Cellulase

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IntEnz Enzyme Nomenclature


Accepted name:
Other names:
9.5 cellulase
β-1,4-endoglucan hydrolase
alkali cellulase
celluase A
cellulase A 3
cellulosin AP
endoglucanase D
pancellase SS
carboxymethyl cellulase
1,4-(1,3;1,4)-β-D-glucan 4-glucanohydrolase
Systematic name:
4-β-D-glucan 4-glucanohydrolase



Will also hydrolyse 1,4-linkages in β-D-glucans also containing 1,3-linkages.

Links to other databases

Enzymes and pathways: NC-IUBMB , BRENDA , ExplorEnz , ENZYME@ExPASy , KEGG , MetaCyc , UniPathway
Protein domains and families: PROSITE:PDOC00511 , PROSITE:PDOC00563 , PROSITE:PDOC00565 , PROSITE:PDOC00640 , PROSITE:PDOC00877 , PROSITE:PDOC51172
Structural data: CSA , EC2PDB
Gene Ontology: GO:0008810
CAS Registry Number: 9012-54-8
UniProtKB/Swiss-Prot: (215) [show] [UniProt]


  1. Datta, P.K., Hanson, K.R. and Whitaker, D.R.
    Improved procedures for preparation and characterization of Myrothecum cellulase. III. Molecular weight, amino acid composition, terminal residues, and other properties.
    Can. J. Biochem. Physiol. 41 : 697-705 (1963). [PMID: 14025219]
  2. Larner, J.
    Other glucosidases.
    In: Boyer, P.D., Lardy, H. and Myrbäck, K. (Eds.) The Enzymes , 2nd ed. vol. 4 , Academic Press , New York , 1960 , 369-378
  3. Myers, F.L. and Northcote, D.H.
    Partial purification and some properties of a cellulase from Helix pomatia.
    Biochem. J. 71 : 749-756 (1959). [PMID: 13651124]
  4. Nishizawa, K. and Hashimoto, Y.
    Cellulose splitting enzymes. VI. Difference in the specificities of cellulase and β-glucosidase from Irpex lacteus.
    Arch. Biochem. Biophys. 81 : 211-222 (1959).
  5. Whitaker, D.R., Hanson, K.R. and Datta, P.K.
    Improved procedures for preparation and characterization of myrothecium cellulase. 2. Purification procedures.
    Can. J. Biochem. Physiol. 41 : 671-696 (1963). [PMID: 1400026]
  6. Hatfield, R. and Nevins, D.J.
    Purification and properties of an endoglucanase isolated from the cell walls of Zea mays seedlings.
    Carbohydr. Res. 148 : 265-278 (1986).
  7. Hatfield, R. and Nevins, D.J.
    Hydrolytic activity and substrate specificity of an endoglucanase from Zea mays seedling cell walls.
    Plant Physiol. 83 : 203-207 (1987). [PMID: 16665203]
  8. Inohue, M., Hayashgi, K. and Nevins, D.J.
    Polypeptide characteristics and immunological properties of exo- and endoglucanases purified from maize coleoptile cell walls.
    J. Plant Physiol. 154 : 334-340 (1999).

[EC created 1961, modified 2001]